• Advanced Courses in Life Sciences

    Header of Evolution at Transmitting Science

Online Course – 2nd Edition

Model-Based Statistical Inference in Evolutionary Biogeography

March 7th-11th, 2022


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This course will be delivered live online

Online live sessions from Monday to Friday from 08:00 to 12:00 (GMT+1, Madrid time zone), plus 10 hours of participants working on their own on assignments exercises between the live sessions.

Course Model-Based Statistical Inference in Ecological and Evolutionary BiogeographyProgram

Check the full Program here


Training in the biological sciences, preferably evolutionary biology. Participants are expected to have a working knowledge of R and must have a personal computer (Windows, Mac, Linux). The use of a webcam and headphones is strongly recommended, as well as a good internet connection.




March 7th-11th, 2022

Schedule and Course length

Online live sessions from Monday to Friday from  08:00 to 12:00 (Madrid time zone).

20 hours of online live lessons, plus 10 hours of participants working on their own on assignments exercises between the live sessions.

30 hours Online.

This course is equivalent to 1 ECTS (European Credit Transfer System) at the Life Science Zurich Graduate School.

The recognition of ECTS by other institutions depends on each university or school.




Places are limited to 18 participants and will be occupied by strict registration order.

Participants who have completed the course will receive a certificate at the end of it.

Nick Matzke instructor for Transmitting Science

Dr. Nick Matzke
University of Auckland
New Zealand

Wallis Bland instructor for Transmitting Science

Wallis Bland
University of Auckland
New Zealand

Haris Saslis coordinator for Transmitting Science

Dr. Haris Saslis
Transmitting Science



  • Intro to R and phylogenies
  • How to read and use phylogenies
  • A short history of historical biogeography methods and assumptions
  • Likelihood-based statistical model choice
  • Phylogenetic biogeography
  • New probabilistic models for historical biogeography in BioGeoBEARS.
  • Using BioGeoBEARS and interpreting results.


  • Biogeographical stochastic mapping.
  • Including geographical and environmental distance in models.
  • Integrating biogeography with traits — trait-dependent dispersal
  • Running analyses over multiPhylo objects (posterior distribution) and interpreting results.
  • State-dependent Speciation/Extinction models (SSE) basics in R
  • State-dependent Speciation/Extinction models (SSE) for large biogeography problems in Julia
  • Integrating GIS & paleogeography data (e.g. GPlates)

 Help session for student projects.



  • Course Fee
  • Early bird (until January 31st, 2022):
  • 520 € *
    (416 € for Ambassador Institutions)
  • Regular (after January 31st, 2022):
  • 590 € *
    (472 € for Ambassador Institutions)
  • The price is VAT inclusive.
    After registration you will receive confirmation of your acceptance on the course. Payment is not required during registration.

You can check the list of Ambassador Institutions. If you want your institution to become a Transmitting Science Ambassador please contact us at communication@transmittingscience.com


Discounts are not cumulative and apply only on the Course Fee. We offer the possibility of paying in two instalments (contact courses@transmittingscience.com).

Former participants will have a 5 % discount on the Course Fee.

20 % discount on the Course Fee is offered for members of some organizations (Ambassador Institutions). If you want to apply to this discount please indicate it in the Registration form (proof will be asked later).

Unemployed scientists living in the country were the course will be held, as well as PhD students based in that country without any grant or scholarship to develop their PhD, could benefit from a 40 % discount on the Course Fee. If you want to ask for this discount, please contact the course coordinator. That would apply for a maximum of 2 places and they will be covered by strict inscription order.